李帥 嚴斌 孟澤洪 楊茂發 楊肖委 周玉鋒



摘要 葉蟬是茶樹上重要的刺吸性害蟲, 常造成茶樹芽葉萎縮不展,甚至焦邊、糊葉。野外調查發現一種特殊的可造成茶樹葉片大面積雪花狀失綠的葉蟬若蟲,采回室內經單頭飼養后獲得其成蟲,并通過成蟲外部形態和雄性外生殖器結構鑒定該害蟲為帕辜小葉蟬Aguriahana paivana (Distant)。本文報道了該害蟲在貴州的發現和為害,首次記述了帕辜小葉蟬若蟲形態特征及其在茶樹上的典型為害狀,并提供了相應的彩色圖片。
關鍵詞 帕辜小葉蟬; 茶樹; 形態特征; 為害狀
中圖分類號: S 433.3
文獻標識碼: A
DOI: 10.16688/j.zwbh.2019171
Identification of Aguriahana paivana (Distant) and preliminary report on its damage to tea plants
LI Shuai1, YAN Bin2, MENG Zehong1*, YANG Maofa2, YANG Xiaowei1, ZHOU Yufeng3*
(1. Guizhou Tea Research Institute, Guizhou Academy of Agricultural Sciences, Guiyang 550006, China;
2. Institute of Entomology of Guizhou University, The Provincial Key Laboratory for Agricultural
Pest Management of Mountainous Region, Guiyang 550025, China; 3. Guizhou Institute
of Biotechnology, Guizhou Academy of Agricultural Sciences, Guiyang 550006, China)
Abstract
The leafhoppers are extremely harmful piercing pest on tea plants and seriously threaten to tea plant (Camellia sinensis) cultivation. It usually leads to bud leaf shrinkage, even causes leaf tissue necrosis. Through field investigation, we found that a special leafhopper could cause mass of snow-like spots on tea leaves with heavy infestation, then we took back the nymph leafhoppers and fed with fresh tea leaves until all of them emerged into adults. Based on the adult morphological features and male genitalia observation, the leafhopper was identified as Aguriahana paivana (Distant). This paper records its occurrence and damage in Guizhou, and firstly describes the morphological features of nymphs and typical infestation characteristics in tea plant. Identification pictures of adults, nymphs and male genitalia were also provided in the paper.
Key words
Aguriahana paivana; tea plant; morphological feature; infestation characteristics
葉蟬是茶樹最重要的刺吸性害蟲類群之一,據統計,在我國為害茶樹的葉蟬種類達60余種[1],其中以小貫小綠葉蟬Empoasca onukii發生為害最為嚴重[2-4],其他諸如茶新折緣葉蟬Neovulturnus testacea[5-6]、大青葉蟬Cicadella viridis[1]、茶扁葉蟬Chanohirata theae[7]等也能造成一定的為害,但絕大多數種類僅限名錄性記載而對其發生為害情況報道甚少。葉蟬類害蟲通常為害茶梢嫩莖嫩葉,造成茶樹芽葉生長遲緩、萎縮不展,嚴重時產生焦邊、糊葉,影響樹勢[7-8]。然而,筆者于2017年底在貴州茶樹害蟲調查過程中卻發現一種造成茶樹葉片產生雪花狀失綠斑點的葉蟬,通過其成蟲外部形態特征和雄性外生殖器解剖觀察,鑒定該害蟲為帕辜小葉蟬Aguriahana paivana。目前,關于帕辜小葉蟬的報道僅見于分類學研究中[9-10],其生物學信息等所知甚少。本文將鑒定結果和為害特征作一報道,記錄該害蟲在貴州的發現和為害,并提供了該蟲成、若蟲的彩色圖片,研究結果可為茶樹害蟲的識別鑒定和科學防治提供參考。
1 材料與方法
1.1 標本采集及飼養
標本采集地位于貴州省黔東南苗族侗族自治州雷公山自然保護區境內,保護區地跨雷山、臺江、劍河、榕江4縣,屬中亞熱帶季風山地濕潤氣候區,年降雨量在1 300~1 600 mm。采集地點海拔1 590 m左右,寄主茶樹位于山地常綠、落葉闊葉混交林下層,周邊雜草灌木叢生,郁閉度較高(圖1a,b)。由于該蟲若蟲喜于葉片背面靜棲刺吸為害,因此,采集時直接以枝剪將攜帶有若蟲的茶樹枝葉剪下置于采樣袋中帶回室內,后將若蟲進行單頭飼養。將以濕棉花包裹葉柄基部的茶樹新鮮葉片置于培養皿(直徑10 cm)內,將若蟲以小毛筆驅趕至葉片上蓋好培養皿蓋,每皿1頭,每日以注射器(5 mL)向棉花上補充水分,并適時更換葉片,直至若蟲羽化為成蟲供鑒定。
通過野外調查及室內飼養觀察發現,帕辜小葉蟬與茶樹重要害蟲脊冠網蝽Stephanitis chinensis Drake, 1948的習性和為害狀極為相似,兩種害蟲均喜聚集葉片背面為害,造成葉片正面呈現細小失綠斑點,而蜜露同為黑色黏稠狀,分布于葉片背面[19],從為害狀上難以區分,但兩種害蟲分屬半翅目Hemiptera葉蟬科Cicadellidae和網蝽科Tingidae,成若蟲形態差異均較大(圖3),若蟲雖體色較為相似,但脊冠網蝽若蟲頭部及腹部背面均具有多根筍狀刺突(圖3e),因而容易區分。
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