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棗實生后代果實性狀變異分析及優(yōu)異種質(zhì)篩選

2025-08-15 00:00:00吉銀中陳有福閆芬芬王玖瑞劉孟軍
果樹學(xué)報 2025年7期
關(guān)鍵詞:果核冬棗母本

中圖分類號:S665.1 文獻標志碼:A 文章編號:1009-9980(2025)07-1407-15

Variation analysis of fruit traits of jujube seedlings and screening of excellentgermplasm

JI Yinzhong',CHENYoufu’,YANFenfen1*,WANGJiurui2,LIUMengjun2

(NationalandocalJonEnneeingLbraofgEfcecyndigualityCultivionndeeprocesngecgf CharacteristiinoiedtrUveitrn; ChinaJujubeResearchCenterofHebeiAgricultural UniversityBaoding07ooo,Hebei,China)

Abstract: 【Objective】 The study aimed to explore the segregation characteristics and genetic variation offruit traits in jujube (Ziziphus jujuba Mill.) seedlings in order to provide a theoretical basis for jujube sedling selection and cultivar breeding. 【Methods】112 jujube seedling trees of Dongzao and Yuhongas were used as research materials,37 traits such as fruit maturity, high yield,appearance quality,sensory quality, nutritional quality and resistance were identified for investigation. The genetic variation of fruit traits were analyzed,and the gray correlation analysis was used to screen excellent jujube sedlings.【Results】The 17 fruit descriptive traits of the two jujube seedling populations segregated a total of 57 types, exhibiting rich variation and a wide range of segregation phenomena. The fruit shape of Dongzao sedling population consisted of 7 types, exhibiting the widest range of segregation, with the flat-round shape resembling that of Dongzao accounting for 40.79% . The fruit shape of Yuhong seedling population exhibited 6 types,with the elongated shape being the most common,comprising

38.89% ,while the conical shape consistent with the Yuhong accounted for 27.78% . Over 44.74% fruits of the two jujube seedling populations was sweet, consistent with Dongzao and Yuhong. Among them, the fruits with thin fruit skin of the Dongzao seedling population accounted for 39.47% , and those with crispy fruit flesh accounted for 57.89% . The variation in single fruit weight was quite significant, and the fruit weight of the Dongzao population ranged from 2.11g to 30.25g. , indicating a broad degree of segregation, nd the fruit weight of 34.21% of the population exceeded the weight of the female parent. Themaximum single fruit weight and fruit diameter were 19.34g and 11.48m ,respectively, higher than those of Dongzao.The average single fruit weight and fruit diameter of Dongzao were both higher than those of the Yuhong population, indicating that Dongzao has a greater potential for selecting large fruit.The coefficient of variation for intrinsic quality traits ranged from 7.51% to 64.72% .Among these, the Dongzao and Yuhong seedling populations showed a rate of exceeding the maternal parent for titratableacid content, vitamin C content,and flavonoid content of over 57.89% .The variation in soluble solids content was considerable, with the Dongzao population exhibiting a rate of exceeding the maternal parent of up to 51.32% ,and the average value was also higher than that of the maternal parent. This indicated that the Dongzao jujube seedlings exhibited a phenomenon of exceeding the maternal parent in soluble solids, which was beneficial for selecting high-soluble solid resources.The maximum values of soluble sugar content in the Dongzao and Yuhong populations were both above 31.73% ,which was advantageous for selecting high-sugar resources. The coeficient of variation for titratable acid content in the two jujube populations was over 47.86% ,indicating a wide variation in titratable acid content. The rate of exceeding the maternal parent for titratable acid in the two offspring reached 66.67% , with the average also higher than that of the maternal parent, facilitating the selection of high-acid resources. The coefficient of variation for vitamin C content in the two offspring was 23.50% ,with the highest variation range of 754.27mg?100g-1 .The ripening stage of Dongzao seedling population ranged from early September to late October.More than 56.58% of the individuals in the seedling populations exhibited good crack resistance and fruiting ability. Based on genetic distribution characteristics,the fruit length and diameter of the Dongzao and Yuhong seedlings both were conformed to a normal distribution. However, the Dongzao population showed a significantly higher occurrence of large single fruit weights compared with the Yuhong population,indicating that the Dongzao offspring possessed greater potential for developing large fruit germplasm. The soluble sugar content and total phenolic content of the Dongzao and Yuhong population were also conformed to a normal distribution. The normal fitting curve for the soluble solids of the Yuhong offspring was longer on the lower side, with a greater concentration of data points on the upper side,indicating that there were more population with higher values of soluble solids. Both the Dongzao and Yuhong population exhibited points outside the upper side of the normal curve for titratable acid,suggesting the presence of high-acid resources in both ofsprings.Notably, the normal fiting curve for vitamin C in the Dongzao population was longer on the upper side and had more data points,indicating that the Dongzao population were likely to produce high vitamin C resources.【Conclusion】A total of 13 excellnt fresh jujube germplasm resources with a comprehensive taste were selected through gray correlation analysis, along with 5 outstanding germplasm resources for dried jujude processing with a drying rate of ≥42.00% . Additionally, 12 specific germplasm resources were identified, including 4 resources with soluble sugar content ?30.00% , 3 resources with titratable acid content ≥1.40% ,and 5 resources with vitamin C content ?591.00mg?100g-1

Key Words: Jujube; Live offspring; Fruit; Variation; Germplasm screening

棗(ZiziphusjujubaMill.)是鼠李科棗屬植物,是中國的傳統(tǒng)果樹之一。棗樹對土壤適應(yīng)性強、耐貧瘠和鹽堿、管理簡約,是經(jīng)濟和生態(tài)兩用型果樹。目前,市場暢銷品種相較單一,而消費者對食品的健康和多樣性需求增加,因此加速棗樹品種改良和新品種選育尤為重要[1-2]。棗實生選種成本低,操作簡單,可以在較短的時間內(nèi)獲得大量的后代,為育種提供高效途徑]。良好的果實品質(zhì)直接影響經(jīng)濟價值,了解果實品質(zhì)等性狀的遺傳規(guī)律,篩選優(yōu)異種質(zhì)資源,對棗品種的遺傳改良和新品種選育極有意義。

果實性狀一直是科技工作者研究的重點之一。對果實性狀的遺傳規(guī)律進行研究,有利于優(yōu)異種質(zhì)資源的篩選。棗樹因花小、去雄困難、坐果率低等特點,傳統(tǒng)的雜交育種存在一定的限制[4。在自然授粉的前提下,利用收集到的母本種子,通過實生的方式對其實生后代果實主要的性狀進行綜合評定,成為了棗樹育種的主要途徑之一[7-9]。目前,對果實品質(zhì)和糖酸組分含量進行研究,發(fā)現(xiàn)后代果實性狀存在廣泛分離[10-14]。以多個棗的實生后代為材料進行研究發(fā)現(xiàn),在多親本的影響下,后代群體數(shù)量性狀出現(xiàn)較為豐富的變異[15-17]。優(yōu)良品種的實生后代群體的單果質(zhì)量均存在廣泛分離,果實糖酸含量變異幅度也較大,有利于高糖、高酸單株的篩選[18]。當前實生后代棗果實性狀研究集中在果實大小、糖酸含量等方面,對不同棗品種實生后代在果實品質(zhì)、成熟期、果核、豐產(chǎn)性及抗性等性狀的系統(tǒng)性和全面性變異研究還有待加強。因此筆者在本研究中以冬棗、雨虹兩個品種的實生后代及其親本為材料,對果實主要性狀進行變異研究,解析不同棗品種實生后代分離變異特征,篩選出特異性較高的優(yōu)株,并通過綜合評價篩選出綜合性狀突出的優(yōu)株,旨在豐富棗種質(zhì)資源,為新品種選育提供理論參考依據(jù)。

1材料和方法

1.1試驗材料

2016年于第一師十團苗木中心試驗基地,采用隔離罩網(wǎng)控制蜜蜂雜交方式,2017年將取得的種仁基質(zhì)播種育苗,定植于塔里木大學(xué)園藝試驗站4號溫室,2021年春季采集長勢健壯實生接穗,嫁接于12團試驗基地,砧木為6年生灰棗樹,株行距為 1m× 2m ,正常田間管理。2023年對結(jié)果性狀穩(wěn)定的112株實生后代進行調(diào)查,其中冬棗實生后代76株、雨虹實生后代36株。

1.2果實成熟期調(diào)查

成熟期:按照全紅期的采樣時間,確定果實成熟的時間,在9月上旬、9月中旬、9月下旬、10月上旬、10月中旬、10月下旬采樣并進行分類。

1.3果實豐產(chǎn)性及抗性調(diào)查

豐產(chǎn)性、裂果率的等級根據(jù)《中國棗種質(zhì)資源》[、段開行等[2]的依據(jù)進行劃分。

1.4 果實性狀的測定

果實外觀品質(zhì):依據(jù)《中國棗種質(zhì)資源》[、《棗種質(zhì)資源描述規(guī)范和數(shù)據(jù)標準》[2],按照標準對果實描述型性狀進行觀察和品嘗,并實時進行拍照記錄,作為參考依據(jù)。

單果質(zhì)量、果實縱橫徑:選取30個發(fā)育正常的棗,用電子天平測定單果質(zhì)量,單位:g,精確到 0.01g 用游標卡尺測定果實縱橫徑,單位: mm ,精確到兩位小數(shù)點。果形指數(shù):果形指數(shù) 果實縱徑/果實橫徑。

果核質(zhì)量:與單果質(zhì)量的材料一致,將棗的果肉剔除后,刷干凈后晾干用電子天平測定,取平均值,單位:g,精確到 0.01g 。

果核指數(shù) 果核縱徑/果核橫徑。

可食率 1%=1 (單果質(zhì)量-果核質(zhì)量)/單果質(zhì)量×100。

果實營養(yǎng)品質(zhì):可溶性固形物含量采用便捷式可溶性固形物測定儀測定,可溶性糖含量采用蒽酮比色法[22測定;可滴定酸含量采用酸堿中和法[22]測定;維生素C含量采用鉬酸銨比色法3測定。可溶性蛋白含量采用[22考馬斯亮藍G-250測定,黃酮含量[24采用硝酸鋁-亞硝酸鈉比色法測定,總酚含量[24]采用福林酚試劑法測定。

1.5優(yōu)異種質(zhì)資源評價篩選方法

利用灰色關(guān)聯(lián)度中的加權(quán)關(guān)聯(lián)法和等權(quán)關(guān)聯(lián)法,10人小組按照鮮食棗綜合評價標準,對兩個棗品種實生后代群體進行綜合評價,按照制干棗綜合評價標準[25-2對雨虹實生后代進行綜合評價,篩選出優(yōu)異的鮮食棗種質(zhì)資源和制干性能優(yōu)異的種質(zhì)資源,同時按照董玉慧2的評價標準進行特異性種質(zhì)資源的篩選,其中綜合口感按照表1進行賦分,制干性能按照表2進行賦分。

1.6數(shù)據(jù)分析

利用Excle2019和SPSS26.0對所有花、果數(shù)據(jù)進行處理,計算其平均值、標準偏差、最小值、最大

表1綜合口感賦分
表2制干性能賦分

值、極差、變異系數(shù)、偏度和峰度等描述型參數(shù),分析其遺傳變異規(guī)律。用Origin2018繪圖。

2 結(jié)果與分析

2.1棗實生后代果實描述型性狀的變異分析

2.1.1果實形狀及顏色性狀分離情況棗實生后代果實形狀及顏色分離情況如表3所示,果實形狀共分離出8種類型,分離廣泛。冬棗后代果實形狀為7種表型,分離范圍最廣,扁圓形后代最多,占比為40.79% ;其次為長圓形的后代,占比為 30.26% 。雨虹后代果形為6種表型,其中長圓形的后代最多,占比為 38.09% ;其次為母本形狀的圓錐形后代,占比為 27.78% 。果實顏色共分離出4種類型,兩個棗品種實生后代群體均以與母本果實顏色一致的紅色后代最多,其中冬棗后代占比為 85.53% 、雨虹后代占比為 91.67% 。

2.1.2果實感官性狀分離情況棗實生后代果實果肉性狀的分離情況如表4所示,6個性狀分離出21種類型,分離廣泛。兩個棗品種實生后代群體均有40% 以上的后代果實風(fēng)味表現(xiàn)為甜,且與母本性狀表現(xiàn)一致。冬棗后代薄果皮占 39.47% ,果肉酥脆占57.89% ,汁液中占 47.37% ,除果肉汁液外,其余性狀均與母本表現(xiàn)一致的后代較多。雨虹后代中有

表3果實形狀及顏色頻率分布情況Table3Fruitshapeandcolorfrequencydistribution
注: *V ”代表母本的對應(yīng)性狀。下同。 Note: σ?σ√ represents the corresponding trait of the female parent.The same below.
表4果實感官性狀的頻率分布情況Table4Thefrequencydistributionoffleshtraits

47.22% 的后代果皮厚,果肉較致密,果肉汁液少,果肉顏色淺綠色,果肉粗細中等,其中果肉質(zhì)地、果肉粗細、果肉顏色、果實風(fēng)味與母本性狀表現(xiàn)一致的后代較多。

2.2 果實大小性狀及可食率的遺傳變異分析

2.2.1果實大小性狀及可食率的分離情況棗實生后代果實大小性狀及可食率分離情況如表5所示,果實大小性狀及可食率分離廣泛,其變異系數(shù)范圍為 1.73%~57.49% ,其中均以冬棗單果質(zhì)量的變異系數(shù)最大,存在較為廣泛的分離,有利于大小果型資源的篩選。可食率變異系數(shù)最小,均在 1.93% 以下,表明可食率的遺傳較為穩(wěn)定,其中冬棗可食率超母本的占 31.58% ,單果質(zhì)量和果實橫徑最大值均出現(xiàn)在冬棗實生后代中,分別高出母本冬棗 19.34g 和 11.48mm 且冬棗后代單果質(zhì)量和果實橫徑平均值均高丁雨虹實生后代群體,表明冬棗實生后代中山現(xiàn)大果資源的潛能相對較高。

表5棗實生后代果實大小及可食率的分離情況Table5Separation of fruit size andedible rate of jujube seedling offspring

2.2.2果實大小性狀的遺傳分布情況對兩個品種實生群體果實大小性狀和可食率進行正態(tài)性檢驗(圖1),冬棗、雨虹實生后代的果實縱徑和果實橫徑均符合正態(tài)分布,果形指數(shù)、可食率不符合正態(tài)分布。其中冬棗后代的單果質(zhì)量正態(tài)擬合曲線上側(cè)尾部較長,單果質(zhì)量數(shù)值點多集中在下側(cè),說明冬棗后代的單果質(zhì)量偏小居多。冬棗和雨虹后代可食率相差不大,但冬棗后代的單果質(zhì)量山現(xiàn)極大果遠多丁雨虹后代,這表明冬棗后代更具備大果型種質(zhì)的創(chuàng)制潛能。在冬棗后代中果形指數(shù)多集中丁1.0附近,說明冬棗后代扁圓形居多。

圖1果實大小及可食率性狀的遺傳分布Fig.1 Geneticdistribution of fruit sizeand edibleratetraits

2.3果實營養(yǎng)品質(zhì)性狀的變異分析

2.3.1果實營養(yǎng)品質(zhì)性狀的分離情況棗實生后代果實內(nèi)在品質(zhì)的分離情況如表6所示,兩個棗群體實生后代的內(nèi)在品質(zhì)性狀均存在較為廣泛的分離,其變異系數(shù)范圍在 7.51%~64.72% 之間。兩個棗實生后代中可溶性固形物含量變異幅度高達 7.79% .且山現(xiàn)超親現(xiàn)象,有利丁高可溶性固形物含量資源的篩選。可溶性糖含量變異幅度高達 16.42% ,存在廣泛分離,冬棗、雨虹后代中可溶性糖含量出現(xiàn)的最高極端值均在 31.73% 以上,有利丁高糖資源的篩選。可滴定酸含量變異系數(shù)均在 47.86% 以上,大于母本的個數(shù)均在 66.67% 以上,其平均值也均高丁母本,山現(xiàn)超親現(xiàn)象,有利丁高酸資源的篩選。維生素C含量的變異系數(shù)均在 23.50% 以上,其最高變異幅度達到 754.27mg?100g-1 ,存在較為廣泛的分離,冬棗維生素C含量大丁母本的占比達到了 100% ,擁有篩選高維生素C含量種質(zhì)資源的優(yōu)勢。黃酮含量的變異系數(shù)均在 50.62% 以上,分離廣泛;冬棗后代總酚含量超過母本的后代占比 85.53% 以上,存在超親現(xiàn)象。蛋白質(zhì)含量在后代中山現(xiàn)超親占比在19.44% 以上。

2.3.2果實營養(yǎng)品質(zhì)性狀的遺傳分布情況對兩個群體內(nèi)在品質(zhì)性狀進行正態(tài)性檢驗,結(jié)果(圖2)表明冬棗、雨虹實生后代可溶性糖含量、總酚含量均符合正態(tài)分布,維生素C含量、可滴定酸含量、黃酮含量不符合正態(tài)分布。雨虹后代可溶性固形物含量的正態(tài)擬合曲線下側(cè)較長,數(shù)量點多集中在上側(cè),表明后代可溶性固形物含量較大的后代較多。冬棗、雨虹后代可滴定酸含量均出現(xiàn)正態(tài)曲線上側(cè)以外的點,這表明兩個后代有高酸資源的山現(xiàn),其中冬棗后代維生素C含量正態(tài)擬合曲線上側(cè)較長,且有數(shù)量點,這表明冬棗后代會山現(xiàn)高維生素C資源。

表6棗實生后代果實營養(yǎng)品質(zhì)的分離情況Table6Jujube seedlingprogeny fruit nutrition quality separation situation
圖2果實營養(yǎng)品質(zhì)遺傳分布Fig.2Genetic distribution of fruit nutritional quality

2.4果核大小性狀的變異分析

2.4.1果核大小性狀的分離情況對兩個棗品種實生后代的果核大小進行測定和分析,結(jié)果如表7所示,果核性狀存在較為廣泛的分離,其變異系數(shù)范圍為 11.74%~63.09% 。冬棗、雨虹實生后代果核質(zhì)量的變異系數(shù)均最大,變異幅度分別為 0.07~1.01g F0.09~0.31g ,其中果核質(zhì)量最小值出現(xiàn)在冬棗實生后代中,且比親本冬棗還少 0.19g ,兩個棗品種實后代群體果核橫徑變異系數(shù)均最小,說明冬棗后代中有更進一步篩選山果大且核小種質(zhì)資源的潛能。

表7棗實生后代果核大小的分離情況Table7 Separationof fruit core size of jujube seedlings

2.4.2果核大小性狀的遺傳分布情況對果核大小進行正態(tài)性檢驗,結(jié)果(圖3)表明冬棗、雨虹實生后代果核縱徑、果核橫徑、核形指數(shù)符合正態(tài)分布,果核質(zhì)量不符合正態(tài)分布。其中冬棗后代果核質(zhì)量的正態(tài)擬合曲線上側(cè)較長,數(shù)量點多集中在曲線下半部分,這表明后代群體果核質(zhì)量、核形指數(shù)性狀數(shù)值偏小的后代較多。

圖3果核性狀的遺傳分布Fig.3 Geneticdistribution ofthekernel traits

2.5果實成熟期、豐產(chǎn)性及抗性遺傳變異分析

2.5.1棗實生后代果實成熟期分離情況對棗實生后代果實的成熟期進行調(diào)查,結(jié)果如表8所示,冬棗實生后代成熟期的分離類型為6種,其中在10月上旬成熟的居多,株數(shù)為24株,占比為 31.58% ;雨虹成熟期的分離類型為3種,其中10月中旬成熟的居多,株數(shù)為22株,占比為 61.11% 。

2.5.2果實果吊比及抗裂果能力分離情況對棗實生后代果吊比及抗裂果能力進行調(diào)查,結(jié)果如表9所示,冬棗、雨虹實生后代果吊比分離類型均為5種,其中冬棗后代果吊比在1以上的后代株數(shù)為43株,占比為 56.57% ;雨虹后代為20株,占比為55.56% ;在抗裂果能力上,冬棗實生后代分離出5種類型,分離范圍廣,其后代裂果率在 20.00% 以下的株數(shù)為73株,占比為 96.06% ;雨虹實生后代分離出3種類型,其后代裂果率在 20.00% 以下的株數(shù)為35株,占比為 97.22% 。

2.6 優(yōu)異種質(zhì)篩選

2.6.1優(yōu)異鮮食種質(zhì)篩選對兩個棗實生后代果實的綜合口感、可滴定酸含量、單果質(zhì)量、可溶性糖含量、果吊比、裂果率、維生素C含量等7個性狀進行綜合評定,通過最終的得分排名篩選出排名靠前綜合能力突出的優(yōu)株,并依據(jù)每個性狀的實際需求和重要性分別給予不同程度的權(quán)重值,依次為0.25、0.1、0.2、0.15、0.1、0.1、0.1 ,依據(jù)加權(quán)關(guān)聯(lián)度,初步選取了排名前13的后代為優(yōu)異鮮食棗種質(zhì)資源(表10)。其他性狀表現(xiàn)如表11所示,13個優(yōu)株口感評價均在75分以上,較為符合鮮食棗的口感需求,其成熟期在9月上旬的有2株、9月中旬的有4株、9月下旬的有3株、10月上旬的有4株,可溶性固形物含量均在35.93% 以上,13株裂果率為 0.00%,S 株果吊比在1以上。

表8棗實生后代果實成熟期的分離情況Table8Separation of fruit maturityof jujube seedlings
表9棗實生后代豐產(chǎn)性及抗裂能力的分離情況Table9Separation of high yield and crack resistance of jujube seedling offspring

2.6.2優(yōu)異制干種質(zhì)篩選對雨虹實生后代果實的制干率、可滴定酸含量、單果質(zhì)量、可溶性糖含量、裂果率、果吊比、果肉質(zhì)地、果肉粗細、果皮厚度9個性狀進行綜合評價,初步篩選出制干性能較高的優(yōu)株,并依據(jù)每個性狀的實際需求和重要性分別給予不同程度的權(quán)重值,依次為 0.20,0.05,0.15,0.10,0.1, 0.1,0.1,0.1,0.1 ,依據(jù)加權(quán)關(guān)聯(lián)度,初步選取了排名前5的后代為優(yōu)異制干棗種質(zhì)資源(表12)。其他性狀表現(xiàn)如表13所示,5個優(yōu)株制干率均在 42.00% 以上,擁有優(yōu)異的抗裂果能力,果吊比均在0.92以上,其中3-2M1、3-2M5、3-2M6果皮厚度和果肉粗細均為滿分。

表10棗實生后代鮮食棗綜合評價Table10 Comprehensive evaluation offresh jujube of jujube seedling offspring
表11優(yōu)異鮮食棗種質(zhì)資源果實性狀Table11 Fruit traitsof excellent fresh jujube germplasmresources
表12棗實生后代制干棗綜合評價Table12Comprehensive evaluation of dried jujube producedbyjujube seedlings
表13優(yōu)異制干棗種質(zhì)資源果實性狀Table13Fruit traits of excellent dried jujube germplasm resources

2.6.3特色種質(zhì)篩選對于棗特色資源的篩選,篩選出可溶性總糖含量大于 30.00% 的高糖資源4份,最高可達 35.33% ;其中冬棗后代2份、雨虹后代2份。可滴定酸含量大于 1.40% 的高酸資源3份,最高可達 1.68% ;其中冬棗后代1份、雨虹后代2份。維生素C含量大于 591.00mg?100g-1 的高維生素C資源5份,最高可達 754.27mg?100g-1; 均為冬棗后代。特色資源主要果實性狀如表14所示。

3討論

果實的外觀和感官品質(zhì)是決定果實經(jīng)濟價值的重要指標[28],對不同實生群體的果實外觀及感官開展遺傳變異比較及性狀分析,有利于篩選果形優(yōu)良、口感豐富的種質(zhì)資源,為實生選種提供參考依據(jù)。筆者在本研究中發(fā)現(xiàn),兩個棗品種實生后代群體果實外觀、感官等性狀出現(xiàn)較大分離,17個性狀共分離出57個類型。冬棗是典型的圓形果實,后代果形出現(xiàn)7種變異,與母本一致的圓形株系分別占9.21% ,其中冬棗后代扁圓形株系最多 (40.79% ,雨虹果實是圓錐形,后代果形出現(xiàn)6種變異,長圓形株系最多 (38.09% ),可見兩個品種實生后代果形變異較豐富。一般在鮮食棗選育中,常常將果肉的汁液量、粗細、質(zhì)地等口感指標作為篩選優(yōu)異鮮食棗的重要依據(jù)[29-31]。陸琦文[17發(fā)現(xiàn)棗實生后代果肉粗細、汁液等性狀有 50% 以上超過母本,出現(xiàn)豐富的變異。本研究中的試驗材料冬棗是果實質(zhì)地酥脆的鮮食品種,實生后代中果肉酥脆的株系分別占 57.89% ,薄果皮的株系分別占 39.47% ,表明冬棗后代在創(chuàng)制口感較好的鮮食棗種質(zhì)方面更具優(yōu)勢。

果實大小是多基因控制的數(shù)量性狀,存在明顯的趨低遺傳傾向[32-34]。對果實大小性狀進行遺傳變異分析和評價,探究不同實生群體的果實大小遺傳變異特點,對大果型實生選種具有重要的指導(dǎo)意義。筆者在本研究中發(fā)現(xiàn),兩個棗實生后代群體果實大小、果核大小均出現(xiàn)廣泛分離,冬棗實生后代單果質(zhì)量超母本株系占比為 34.21% ,果實橫徑超母本株系占比為 38.16% ,最終呈現(xiàn)冬棗后代扁圓形株系較多,同時冬棗實生后代單果質(zhì)量出現(xiàn)較為極端的變異株型,這進一步表明冬棗實生后代在果型種質(zhì)的篩選方面更具潛力。李鵬麗3在棗實生后代研究中發(fā)現(xiàn),果實的縱徑、橫徑變異最大,變異系數(shù)分別為 28.42% 和 21.39% ,相比本研究中最高果實縱徑、橫徑變異系數(shù)要低11.58和12.82百分點,這可能是實生群體不同所導(dǎo)致。兩個實生后代群體可食率變異范圍在 0.89%~1.93% ,變異程度相對較小,袁野等[15]在蜂蜜罐實生后代中可食率變異為 0.05% ,相比本研究中最低可食率變異要低0.84百分點,這可能是不同授粉品種所影響的結(jié)果。兩個群體比較,冬棗實生群體單果質(zhì)量超親率達 34.21% ,雨虹后代可溶性糖含量超親率 25.00% 為最高。

表14特色資源主要果實性狀Table14Mainfruittraitsofcharacteristicresources

實生后代株系雜合度較高,其果實內(nèi)在品質(zhì)性狀會出現(xiàn)較為廣泛的分離,這有利于特異性種質(zhì)資源的篩選[3。筆者在本研究中發(fā)現(xiàn),兩個棗品種群體實生后代內(nèi)在品質(zhì)變異范圍在 7.51%~75.33% 之間,分離廣泛,維生素C是人體不可或缺的營養(yǎng)元素,可預(yù)防敗血癥,筆者在本研究中篩選出的5份高維生素C資源含量均在 593.25mg?100g-1 以上,最高可達 754.27mg?100g-1 。相比董玉慧 [27]248 棗品種維生素C平均含量為 414.00mg?100g-1 而言高出179.25mg?100g-1 。高糖高酸的果實有利于加工口感豐富的特色果干和果汁,筆者在本研究中所獲得的3份高酸資源含量在 1.41% 以上,相比董玉慧[27]248棗品種最高可滴定酸 1.02% 高出0.39百分點,最高可達 1.68% ,所獲得的7份高糖資源含量在30.24% 以上,相比董玉慧 [27]248 棗品種平均可溶性糖26.0% 高出4.24百分點,最高可達 35.33% 。綜合分析,果實質(zhì)地酥脆、汁液中等的種質(zhì)可作為鮮食棗選育的候選材料,制干率高、果肉厚且口感軟糯的種質(zhì)可進一步挖掘制干優(yōu)系,高糖、高酸、高維生素C等特異優(yōu)系可進一步為功能食品開發(fā)等深加工品種選育提供候選材料,從而加快棗新優(yōu)特品種的選育進程。

4結(jié)論

棗兩個品種實生后代群體果實性狀出現(xiàn)豐富變異, 40% 左右后代果實與母本果形相似。冬棗實生后代果肉酥脆的株系占比高達 57.89% ,出現(xiàn)大果、扁圓形株系較多。兩個群體可滴定酸含量、維生素C含量、黃酮含量超母本率超過 57.89% ,其中冬棗后代出現(xiàn)高果實維生素C含量和可滴定酸含量極端株系,56.58% 以上的后代具備較強的抗裂果能力和較強的坐果能力。雨虹后代果實可溶性糖含量變異廣泛,出現(xiàn)了可溶性固形物含量和高可滴定酸含量極端株系。初步篩選出綜合性狀優(yōu)異的鮮食棗種質(zhì)13份;綜合性狀優(yōu)異的制干棗種質(zhì)5份;特異種質(zhì)12份,其中高糖種質(zhì)4份;高酸種質(zhì)3份;高維生素C種質(zhì)5份。

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